Positive planes - one has to fill the copper area with vectors (traces) or in some packages, areas. When it converted to Gerber files, most of the packages will convert filled areas into smaller vectors and it will result in increased Gerber file size.
If negative layers are used, this can be avoided and also it is easier to draw the split planes in negative as we need to draw the vectors comprising the devided area between two planes.
First file is a positive plane (top layer)
Second file is a Negative plane (Internal layer2 of a 4 layer board)
On the positive planes, the area surronding the traces and pads are etched away from the copper layer.
On the Negative plane, what shows are pads and thermal reliefs are etched out of the copper, so the black area on this picture is actually solid copper.
I've also done Positive layers on internal layers where I need to put traces and copper areas in the layout - that would look the same as the Top layer pic.
And, traces can also be embedded in Negative layers, where the net/trace would have an enclosed outline around it to create a mote of conductivity.